1Department of Veterinary Infectious Diseases, College of Veterinary Medicine, Chonnam National University, Gwangju 61186, Korea 2Biology Department at Morrissey College of Arts and Sciences, Boston College, Chestnut Hill, MA 02467, USA 3Department of Pulmonary Immunology, Center for Pulmonary and Infectious Diseases Control, University of Texas Health Science Center at Tyler, Tyler, Texas 75708, USA